Аннотация:The elaborated Thylakoid Model (TM) allowed fitting to both FI and ΔA810 data monitored (at PFD 200 μmol photons m2 s1) on dark-adapted pea leaves during the 20 s time range. The two-wave kinetics of O(JI)PSMT and OABCD were reproduced in their parallel and antiparallel phases with the parameter quantification concerned to electron and proton transfer in thylakoid membrane, lumen, stroma compartments. (t) and pHL/S(t) dynamics was found physiologically relevant while the TM fitted FI and ΔA810 data under assumption that 20% of F0 is attributed to PSI. Changing in the model the buffer capacity from 43 to 81 mM gave the peak (t) increasing from 8 to 15 mV at reasonable ranges of H, K, Cl fluxes. The previous TM simulations that assumed the unvaried activation state of FNR and stromal enzyme were refined. We introduced the time-dependent rate constant to model the NADP reduction by ferredoxin supposing the FNR activation effect in the time domain of 20–40 s after the start of a low intensity illumination. Transitions between the linear and cyclic modes, concurrent with ATPase operation can be clarified over a wide time-scale due to quantitative analysis of the light induction stages preceding the stationary state.